Great Britain's Jody Cundy saw his 20-year reign as para cycling's C4 kilo time trial world champion come to an end at the para cycling track world championships in the Netherlands.
The 48-year-old, who won his first world title in 2006 and had held the rainbow jersey through 15 consecutive championships, finished with a time of one minute and 4.93 seconds to claim silver.
Australia's Korey Boddington took gold, beating Cundy's time by 2.25 seconds on the final run.
Cundy, a nine-time Paralympic champion with three medals in para swimming between 1996 and 2000, had acknowledged before the event that "age was catching up with him" and that adding to his 23 track cycling world titles would be "hard."
Elsewhere on the opening day, Great Britain's Rebecca Newark won gold in the women's C5 kilo time trial. In the women's B kilo time trial, Sophie Unwin and pilot Jenny Holl secured silver, while Elizabeth Jordan and pilot Anna Whitworth Hay took bronze.
In the men's 4km pursuit, the British pair Morgan Fice-Noyes and Christopher Latham earned silver, and Finlay Graham added bronze in the men's C3 sprint final.
